List of pterosaur genera

This list of pterosaurs is a comprehensive listing of all genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (nomen dubium), or were not formally published (nomen nudum), as well as junior synonyms of more established names, and genera that are no longer considered pterosaurian. The list currently includes 205 genera.

Scope and terminology

There is no official, canonical list of pterosaur genera, but the most thorough attempts can be found at the Pterosauria section of Mikko Haaramo's Phylogeny Archive, the Genus Index at Mike Hanson's The Pterosauria, supplemented by the Pterosaur Species List, and in the fourth supplement of Donald F. Glut's Dinosaurs: The Encyclopedia series.

Authors and year

The authors column lists the authors of the formal description responsible for the erection of the genus listed. They are not necessarily the same as the authors of the type species as sometimes a species from one genus is determined sufficiently distinct to warrant the erection of a new genus to house it. If this is the case, only the latter authors will be listed. The year column notes the year the description was published.

Status

Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:

• Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.
• Nomen nudum (Latin for "naked name"): A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. If the name is later formally published, that name is no longer a nomen nudum and will be italicized on this list. Often, the formally published name will differ from any nomina nuda that describe the same specimen.
• Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid (as are all subsequent uses) and the name must be replaced. As preoccupied names are not valid generic names, they will also go unitalicized on this list.
• Nomen dubium (Latin for "dubious name"): A name describing a fossil with no unique diagnostic features. As this can be an extremely subjective and controversial designation, this term is not used on this list.

Age

The age column denotes the epoch of geologic time to which the fossils date. Genera that are invalid, misidentified, or otherwise do not represent a valid pterosaur are listed as age N/A because there was never a time in which a pterosaur by that generic name actually lived.

Location and notes

The location column designates the geographic region where remains of the relevant genus have been found. The regions used are continents except in the case of smaller landmasses (e.g. Cuba). Political bodies, being non-existent in the Mesozoic are not used to indicate genera locations. Genera that are invalid, misidentified, or otherwise do not represent a valid pterosaur are listed as location N/A because there was never a place in which a pterosaur by that generic name actually lived. The notes column is a collection of annotations on the scientific significance and taxonomic history of listed genera, as well as elaborations on the information presented in other columns.

The list

Genera

Genus Authors Year Status Age Location Notes Aerodactylus Vidovic and Martill 2014 Valid Late Jurassic Europe Aerotitan Novas et al. 2012 Valid. Late Cretaceous S. America Aetodactylus Myers 2010 Valid. Late Cretaceous N. America Aidachar Nesov 1981 Misidentification. N/A N/A Actually a teleost fish whose remains were originally mistaken for jaw fragments of a ctenochasmatid. The mistake was corrected in 1986. Alamodactylus Andres Myers 2013 Valid. Late Cretaceous N. America Alanqa Ibrahim et al. 2010 Valid Late Cretaceous Africa Amblydectes Hooley 1914 Nomen dubium. Early Cretaceous Europe Angustinaripterus He Xinlu et al. 1983 Valid. Middle Jurassic Asia Known from a single skull recovered from the Dashanpu Formation. Its Dorygnathus-like teeth suggest it had a piscivorous diet. Anhanguera Campos Kellner 1985 Valid. Early Cretaceous S. America A piscivorous ornithocheirid from the Santana Formation. A study of its anatomy helped resolve controversy regarding pterodacyloids' posture while on the ground. Anurognathus Döderlein 1923 Valid. Late Jurassic. Europe. A tiny (50 cm wingspan) insectivore known only from two skeletons recovered from the Solnhofen Formation. Apatomerus Williston 1903 Misidentification. Lower Cretaceous N. America Probably a misidentified plesiosaur. Arambourgiania Nesov Kanznyshkina Cherepanov 1987 Valid. Late Cretaceous Asia Aralazhdarcho Averianov 2007 Valid. Late Cretaceous Asia Araripedactylus Wellnhofer 1977 Valid. Early Cretaceous S. America Araripesaurus Price 1971 Valid. Early Cretaceous S. America Archaeoistiodactylus Lü Fucha 2011 Valid. Middle Jurassic Asia Ardeadactylus Bennett 2013 Valid. Late Jurassic Europe Arthurdactylus Frey Martill 1994 Valid. Early Cretaceous S. America Aurorazhdarcho Frey Meyer Tischlinger 2011 Valid. Late Jurassic. Europe. Aussiedraco Kellner Rodrigues Costa 2011 Valid. Early Cretaceous Australia Austriadactylus Dalla Vecchia Wild Reitner 2002 Valid. Upper Triassic Europe Avgodectes Peters 2004 Jr. synonym. N/A N/A Probable junior synonym of Haopterus. Azhdarcho Nesov 1984 Valid. Late Cretaceous Asia Bakonydraco Weishampel Jianu 2005 Valid. Late Cretaceous Europe Banguela Headden Campos In press Valid. Early Cretaceous S. America Barbosania Elgin Frey 2011 Valid. Late Cretaceous S. America Batrachognathus Rjabinin 1948 Valid. Late Jurassic. Asia. Beipiaopterus J.-C. Lü 2003 Valid. Early Cretaceous Asia Belonochasma Broili 1939 Misidentification. N/A N/A A non-pterosaurian gnathostome. Bellubrunnus Hone et al. 2012 Valid. Late Jurassic Europe Bennettazhia Nesov 1991 Valid. Early Cretaceous N. America Bogolubovia Nesov A. A. Yarkov 1989 Valid. Late Cretaceous Asia Boreopterus Lü Qiang 2005 Valid. Early Cretaceous Asia Brachytrachelus Giebel 1852 Preoccupied. N/A N/A preoccupied name; now known as Scaphognathus Brasileodactylus Kellner 1984 Valid. Early Cretaceous S. America Cacibupteryx Gasparini Fernández de la Fuente 2004 Valid. Late Jurassic Cuba Caiuajara Manzig et al. 2014 Valid. Late Cretaceous S. America Camposipterus Rodrigues Kellner 2013 Valid. Early Cretaceous Europe "Campylognathus" Plieninger 1894 Preoccupied. N/A N/A preoccupied name; now known as Campylognathoides Campylognathoides Strand 1928 Valid. Early Jurassic Europe. Carniadactylus Dalla Vecchia 2009 Valid. Late Triassic Europe Formerly Eudimorphodon rosenfeldi. Cathayopterus Wang Zhonghe 2006 Valid. Early Cretaceous Asia Caulkicephalus Steel Martill et al. 2005 Valid. Early Cretaceous Europe Caupedactylus Kellner 2013 Valid. Early Cretaceous S. America Caviramus Fröbisch Fröbisch 2006 Valid. Upper Triassic Europe Cearadactylus Leonardi Borgomanero 1985 Valid. Early Cretaceous S. America Changchengopterus Lü 2009 Valid. Late Jurassic Asia Chaoyangopterus Wang Zhou 2003 Valid. Early Cretaceous Asia Cimoliopterus Rodrigues Kellner 2013 Valid. Late Cretaceous Europe Cimoliornis Owen 1846 Nomen nudum. N/A N/A Coloborhynchus Owen 1874 Valid. Early Cretaceous Europe N. America S. America Comodactylus Galton 1981 Valid. Late Jurassic N. America Cretornis Fritsch 1880 Nomen dubium. Late Cretaceous Europe Criorhynchus Owen 1874 Jr. synonym. N/A N/A Jr. synonym of Ornithocheirus. Ctenochasma von Meyer 1852 Valid. Late Jurassic. Europe. Cuspicephalus Martill Etches 2013 Valid. Late Jurassic Europe Cycnorhamphus Seeley 1870 Valid. Late Jurassic. Europe. "Daitingopterus" Maisch Matzke Ge Sun 2004 Nomen nudum. Late Jurassic Europe Darwinopterus Lü Unwin et al. 2009 Valid. Middle Jurassic Asia Intermediate form between rhamphorhynchoids and pterodactyloids. Dawndraco Kellner 2010 Valid. Late Cretaceous N. America Dendrorhynchoides S.-A. Ji Q. Ji Padian 1999 Valid. Late Jurassic or Early Cretaceous Asia "Dendrorhynchus" S.-A. Ji Q. Ji 1998 Preoccupied. N/A N/A Preoccupied name; now known as Dendrorhynchoides. Dermodactylus Marsh 1881 Valid. Late Jurassic N. America Dimorphodon Owen 1859 Valid. Early Jurassic Europe. Diopecephalus Seeley 1871 Jr. synonym. N/A N/A Jr. synonym of Pterodactylus. Domeykodactylus Martill Frey et al. 2000 Valid. Early Cretaceous S. America Doratorhynchus Seeley 1875 Nomen vanum Late Jurassic or Early Cretaceous Europe Dorygnathus Wagner 1860 Valid. Early Jurassic Europe. Dsungaripterus Young 1964 Valid. Late Jurassic-Early Cretaceous Africa. Asia. Elanodactylus Andres Ji, Q. 2008 Valid. Early Cretaceous Asia Eoazhdarcho Lü Qiang 2005 Valid. Early Cretaceous Asia Eopteranodon Lü, J.C. Zhang 2005 Valid. Early Cretaceous Asia Eosipterus Ji Ji 1997 Valid. Early Cretaceous Asia Eudimorphodon Zambelli 1973 Valid. Late Triassic Europe. Europejara Vullo et al. 2012 Valid. Early Cretaceous Europe Eurazhdarcho Vremir et al. 2013 Valid. Late Cretaceous Europe Eurolimnornis Kessler Jurcsák 1986 Valid. Early Cretaceous Europe Originally described as a bird, subsequently reinterpreted as a pterosaur. Faxinalipterus Bonaparte, et al. 2010 Valid. Late Triassic S. America Feilongus Wang, Kellner, et al. 2005 Valid. Early Cretaceous Asia Fenghuangopterus Lü Fucha Chen 2010 Valid. Middle Jurassic Asia Gallodactylus Fabre 1974 Jr. synonym. N/A N/A Jr. synonym of Cycnorhamphus. Geosternbergia Miller 1978 Jr. synonym. N/A N/A Jr. synonym of Pteranodon. Gegepterus X. Wang Kellner et al. 2007 Valid. Early Cretaceous Asia Germanodactylus Yang 1964 Valid. Late Jurassic. Europe. Gladocephaloideus Lü Ji Wei Liu 2011 Valid. Early Cretaceous Asia Gnathosaurus von Meyer 1833 Valid. Late Jurassic Europe Guidraco Wang et al. 2012 Valid. Early Cretaceous Asia Gwawinapterus Arbour Philip J. Currie 2011 Misidentification. Late Cretaceous N. America Initially thought to be an istiodactylid pterosaur, but subsequently reinterpreted as an indeterminate saurodontid fish. Hamipterus Wang et al. 2014 Valid. Early Cretaceous Asia Haopterus Wang X. Lü J. 2001 Valid. Early Cretaceous Asia Harpactognathus Carpenter Unwin et al. 2003 Valid. Late Jurassic N. America Hatzegopteryx Buffetaut Grigorescu Csiki 2002 Valid. Late Cretaceous Europe Herbstosaurus Casamiquela 1974 Valid. Late Jurassic S. America Hongshanopterus X. Wang Campos et al. 2008 Valid. Early Cretaceous Asia Huanhepterus Dong 1982 Valid. Late Jurassic Asia Huaxiapterus J. Lü C. Yuan 2005 Valid. Early Cretaceous Asia Ikrandraco Wang et al. 2014 Valid. Early Cretaceous Asia Ingridia Unwin Martill 2007 Jr. synonym. N/A N/A Objective Jr. synonym of Tupandactylus. Istiodactylus Howse Milner Martill 2001 Valid. Early Cretaceous Europe Jeholopterus X. Wang Z. Zhou 2002 Valid. Middle Jurassic or Early Cretaceous Asia Jianchangnathus Cheng Wang Jiang Kellner 2012 Valid. Middle Jurassic Asia Jianchangopterus Lü Bo 2011 Valid. Middle Jurassic Asia Jidapterus Dong Sun Wu 2003 Valid. Early Cretaceous Asia Kepodactylus Harris Carpenter 1996 Valid. Late Jurassic N. America Kunpengopterus X. Wang Kellner et al. 2010 Valid. Middle Jurassic Asia Kryptodrakon Andres Clark Xu 2014 Valid. Middle Jurassic and/or Late Jurassic Asia Lacusovagus Witton 2008 Valid. Early Cretaceous S. America Laopteryx Marsh 1881 Valid. Late Jurassic N. America Liaoningopterus X.-L. Wang Z.-H. Zhou 2003 Valid. Early Cretaceous Asia Liaoxipterus Z. Dong J. Lü 2005 Valid. Early Cretaceous Asia Limnornis Kessler Jurcsák 1984 Preoccupied. Early Cretaceous Europe Originally described as a bird; fossils later renamed Palaeolimnornis. Linlongopterus Rodrigues et alii 2015 Valid. Early Cretaceous Asia "Lithosteornis" Gervais 1844 Nomen nudum. N/A N/A Nomen nudum. Lonchodectes Hooley 1914 Valid. Early Cretaceous to Late Cretaceous Europe Lonchodraco Rodrigues Kellner 2013 Valid. Early Cretaceous to Late Cretaceous Europe Lonchognathosaurus Maisch Matzke Ge Sun 2004 Valid. Early Cretaceous Asia Longchengpterus L. Wang L. Li et al. 2006 Valid. Early Cretaceous Asia Ludodactylus Frey Martill Buchy 2003 Valid. Early Cretaceous S. America Maaradactylus Bantim et alii 2014 Valid Early Cretaceous S. America Macrotrachelus Giebel 1852 Jr. synonym. N/A N/A Jr. synonym of Pterodactylus. Mesadactylus Jensen Padian 1989 Valid. Late Jurassic N. America Microtuban Elgin Frey 2011 Valid. Late Cretaceous Africa Moganopterus Lü et al. 2012 Valid. Early Cretaceous Asia Montanazhdarcho Padian Horner de Ricqlès 1993 Valid. Late Cretaceous N. America Muzquizopteryx Frey Buchy et al. 2006 Valid. Late Cretaceous N. America Mythunga Molnar Thulborn 2008 Valid. Early Cretaceous Australia Navajodactylus Sullivan Fowler 2011 Valid. Late Cretaceous N. America Nemicolopterus X. Wang Kellner et al. 2008 Valid. Early Cretaceous Asia Nesodactylus Colbert 1969 Valid. Late Jurassic Cuba Nesodon Jensen Ostrom 1977 Lapsus calami. N/A N/A Misspelling of Nesodactylus, also preoccupied by a toxodont. Ningchengopterus J. Lu 2009 Valid. Early Cretaceous Asia Noripterus Yang 1973 Valid. Early Cretaceous Asia Normannognathus Buffetaut J.-J. Lepage G. Lepage 1998 Valid. Late Jurassic Europe Nurhachius Wang Kellner et al. 2003 Valid. Early Cretaceous Asia Nyctodactylus Marsh 1881 Jr. synonym. N/A N/A Jr. synonym of Nyctosaurus. Nyctosaurus Marsh 1876 Valid. Late Cretaceous N. America. S. America. "Odontorhynchus" Stolley 1936 Jr. synonym. N/A N/A Jr. synonym of Rhamphorhynchus. "Oolithorhynchus" Whalley 2000 Nomen nudum. Manuscript name about which almost nothing is known. Ornithocephalus von Sömmering 1812 Jr. synonym. N/A N/A Jr. synonym of Pterodactylus. Ornithocheirus Seeley 1869 Valid. Late Jurassic. Africa. Australia. Europe. S. America. Ornithodesmus Seeley 1887 Misidentification. N/A N/A Misidentified dromaeosaurid. Ornithopterus Fitzinger 1843 Jr. synonym. N/A N/A Jr. synonym of Rhamphorhynchus. Ornithostoma Seeley 1871 Valid. Late Cretaceous Europe "Osteornis" Gervais 1844 Nomen nudum. N/A N/A Nomen nudum, but appeared to be a Jr. synonym of Ornithocheirus anyway. Pachyrhamphus Fitzinger 1843 Preoccupied. N/A N/A preoccupied name; now known as Scaphognathus Palaeocursornis Kessler Jurcsák 1986 Valid. Early Cretaceous Europe Originally described under the name Limnornis as a bird; Limnornis was preoccupied, and the fossils were subsequently reinterpreted as pterosaurian. Palaeornis Mantell 1844 Preoccupied. N/A N/A preoccupied name. Paranurognathus Peters 2005 Jr. synonym. N/A N/A Jr. synonym of Anurognathus. Parapsicephalus von Arthaber 1919 Valid. Early Jurassic Europe Peteinosaurus Wild 1978 Valid. Late Triassic Europe. "Phobetor" Bakhurina 1986 Jr. synonym. N/A N/A Jr. synonym of Noripterus. Phosphatodraco Pereda-Suberbiola Bardet et al. 2003 Valid. Late Cretaceous Africa Piksi Varricchio 2002 Valid. Late Cretaceous N. America Originally described as a bird, subsequently reinterpreted as a pterosaur. Plataleorhynchus Howse Milner 1995 Valid. Late Jurassic to Early Cretaceous Europe Prejanopterus Vidarte Calvo 2010 Valid. Early Cretaceous Europe Preondactylus Wild 1983 Valid. Late Triassic Europe "Pricesaurus" Bonaparte Sanchez 1986 Nomen nudum. Early Cretaceous S. America Procoelosaurus Atanassov 2002 Nomen ex dissertatione. Ptenodactylus Seeley 1869 Preoccupied N/A N/A Preoccupied name. Ptenodracon Lydekker 1888 Jr. synonym N/A N/A Junior synonym of Ctenochasma. Pteranodon Marsh 1876 Valid. Late Cretaceous N. America. Ptéro-dactyle Cuvier 1809 N/A N/A Renamed Pterodactylus. Pterodactylus Rafinesque 1815 Valid. Late Jurassic. Africa. Europe. Pterodaustro Bonaparte 1970 Valid. Early Cretaceous S. America. Pterofiltrus Jiang Wang 2011 Valid. Early Cretaceous Asia Pteromimus Atanassov 2002 Nomen ex dissertatione. Pteromonodactylus Teryaev 1967 Jr. synonym. N/A N/A Jr. synonym of Rhamphorhynchus. Pterorhynchus Czerkas Q. Ji 2002 Valid. Late Jurassic Asia Pterotherium Fischer von Waldheim 1813 Jr. synonym. N/A N/A Jr. synonym of Pterodactylus. Puntanipterus Bonaparte Sanchez 1975 Valid. Late Jurassic to Early Cretaceous S. America Qinglongopterus J. Lü Unwin et al. 2012 Valid. Middle Jurassic Asia Quetzalcoatlus Lawson 1975 Valid. Late Cretaceous N. America. Radiodactylus Andres Myers 2013 Valid. Early Cretaceous N. America Raeticodactylus Stecher 2008 Jr. synonym N/A N/A Jr. synonym of Caviramus Rhabdopelix Cope 1870 Misidentification. N/A N/A At first it was thought to be a Triassic pterosaur, but is now known to be (at least in part) a kuehneosaurid. Rhamphinion Padian 1984 Valid. Early Jurassic N. America Rhamphocephalus Seeley 1880 Valid. midjur Europe Rhamphorhynchus von Meyer 1846 Valid. Late Jurassic. Africa. Europe. Santanadactylus de Buisonjé 1980 Valid. Early Cretaceous S. America Scaphognathus Wagner 1861 Valid. Late Jurassic. Europe. Sericipterus Andres Clark X. Xing 2010 Valid. Late Jurassic Asia Shenzhoupterus J. Lü Unwin et al. 2008 Valid. Early Cretaceous Asia Sinopterus X. Wang Z. Zhou 2003 Valid. Early Cretaceous Asia Siroccopteryx Mader Kellner 1999 Jr. synonym. N/A N/A Jr. synonym of Coloborhynchus. Sordes Sharov 1971 Valid. Late Jurassic. Asia. Sultanuvaisia Nesov 1981 Misidentification. Late Cretaceous Asia. Actually a fish. Tapejara Kellner 1989 Valid. Early Cretaceous S. America Tendaguripterus Unwin Heinrich 1999 Valid. Late Jurassic Africa Thalassodromeus Kellner Campos 2002 Valid. Early Cretaceous S. America "Titanopteryx" Arambourg 1959 Preoccupied. Late Cretaceous Asia Preoccupied by a simuliid blackfly. It was later renamed Arambourgiania. Tribelesodon Bassani 1886 Misidentification. N/A N/A At first it was thought to be a Triassic pterosaur but is now known to be a misinterpreted specimen of the prolacertiform Tanystropheus. Tropeognathus Wellnhofer 1987 Valid. Early Cretaceous S. America One of the largest Ornithocheirids. Tupandactylus Kellner Campos 2007 Valid. Early Cretaceous S. America Tupuxuara Kellner Campos 1988 Valid. Late Cretaceous N. America S. America Uktenadactylus Rodrigues Kellner 2008 Disputed. Early Cretaceous N. America Unwindia Martill 2011 Valid. Late Cretaceous S. America Utahdactylus Czerkas Mickelson 2002 Disputed. N/A N/A Probably a non-pterosaurian reptile. Vectidraco Naish Simpson Dyke 2013 Valid. Early Cretaceous Europe Volgadraco Averianov Arkhangelsky Pervushov 2008 Valid. Late Cretaceous. Asia. Wenupteryx Codorniú Gasparini 2013 Valid. Late Jurassic S. America Wukongopterus X. Wang Kellner et al. 2009 Valid. Middle Jurassic Asia "Wyomingopteryx" Bakker 1994 Nomen nudum. Nomen nudum. Yixianopterus J. Lü S. Ji et al. 2006 Valid. Early Cretaceous Asia Appears to be valid, but little is known Zhejiangopterus Z. Cai F. Wei 1994 Valid. Late Cretaceous Asia Zhenyuanopterus J. Lü 2010 Valid. Early Cretaceous Asia		File:Angustinaripterus NT.jpg Angustinaripterus File:Anhanguera.jpg Anhanguera Anurognathus File:Pterodactylus antiquus 01 by Line1.jpg Aurorazhdarcho Bakonydraco File:Campylogn DB.jpg Campylognathoides File:Cearadactylus.jpg Cearadactylus Coloborhynchus File:Ctenochasma elegans-BackgroundKnockedOut-ROM-Dec29-07.png Ctenochasma Dimorphodon Dorygnathus File:Dsungaripterus weii.jpg Dsungaripterus Eudimorphodon File:Feilongus NT.jpg Feilongus File:Germ rhamph DB.jpg Germanodactylus File:HoopterusGracilis-PaleozoologicalMuseumOfChina-May23-08.jpg Haopterus File:Hatzegopteryx BW.jpg Hatzegopteryx File:IstiodactylusDB3.jpg Istiodactylus File:Jeholopterus ningchengensis.png Jeholopterus Liaoningopterus File:Nemicolopterus.jpg Nemicolopterus File:Ningchengopterus BW.jpg Ningchengopterus File:Phobetor parvis.jpg Noripterus File:Nurhachius NT.jpg Nurhachius File:Nyctosaurus DB.jpg Nyctosaurus File:Ornithocheirus BW.jpg Ornithocheirus File:Preondactylus apatomerus.jpg Preondactylus Pteranodon Pterodactylus Error creating thumbnail: File missing Pterodaustro Quetzalcoatlus Rhamphorhynchus File:ScaphognDB.jpg Scaphognathus Sordes File:Tapejara.jpg Tapejara File:Thalassodromeuswitton2.jpg Thalassodromeus File:TapimpDB.jpg Tupandactylus Tupuxuara Zhejiangopterus

Ichnogenera

Genus Authors Year Status Age Location Notes Agadirichnus Ambroggi Lapparent 1954 Disputed. uK Africa Haenamichnus Hwang Huh et al. 2002 Valid. uK Asia. More than 5 times as large as Pteraichnus, these tracks were probably made by azhdarchids. Kouphichnium Nopcsa 1923 Misidentification. N/A N/A The trackmaker was probably a limulid. Pteraichnus Stokes 1957 Valid. uJ Asia Europe N. America Possible junior synonym of Agadirichnus. Some tracks attributed to Pteraichnus have been considered crocodilian in origin by some scientists (Padian, 1984) however, recent studies in the past decade have overturned Padian's thoughts. Purbeckopus Delair 1963 Nomen dubium. lK Europe.		Haenamichnus

Oogenera

Although pterosaur eggs are known, some with complete embryos, no oogenera have been erected to house them. The holotype of the oospecies Oolithes sphaericus was briefly considered by Harry Govier Seeley to be pterosaurian in origin, although this attribution was dismissed before the formal erection of that oogenus.

See also

• List of dinosaurs
• List of plesiosaurs
• List of pterosaur classifications
• Pterosaur

External links

• Pterosaur FAQs, by Raymond Thaddeus C. Ancog.
• The Pterosaur Database, by Paul Pursglove.
• Mark Witton's Pterosaur Art